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Edward Schilling


Research Interest

Plant systematics


1978 - Ph.D., Indiana University


My research interests are in higher plant systematics with a focus on members of the sunflower family (Asteraceae).

Molecular Phylogenetic Analysis of Eupatorieae

A collaborative study being carried out together with Dr. Jose L. Panero of the University of Texas involves a systematic analysis of tribe Eupatorieae. This large group is primarily restricted to the New World, where it has produced an amazing amount of species-level diversity. Molecular phylogenetic analyses are being carried out with the goal of resolving issues of classification, and assessing the morphological changes associated with the success of this group.

Systematic Studies of Helianthus

The genus Helianthus comprises approximately 50 species native  to North America, and includes the agronomically important common sunflower, Helianthus annuus. Currently the focus of my studies are on the perennial species of the genus, with a particular emphasis on resolving issues of classification and working out the relationships of some of the polyploids, such as H. hirsutus, H. strumosus, and H. tuberosus.

Other Research in my Laboratory

Systematic and Evolutionary Studies of Trillium - Trillium and its relatives of tribe Parideae are long-lived spring-blooming wildflowers that exhibit some of the largest genome sizes documented for plants.  We are conducting systematic and evolutionary studies of the group to work out its alpha taxonomy and basic classification, as well as to address synthetic questions regarding the biogeography and evolution of the group.


Visit Google Scholar for more publications.

  • Schilling, E. E. and K. S. Grubbs. 2016. Systematics of the Eupatorium mohrii complex (Asteraceae). Systematic Botany. DOI 10.1600/036364416X692361
  • Rivera, Vanessa L., J. L. Panero, Schilling, E. E., B. S. Crozier, and Marta D. Moraes. 2016. Origins and recent radiation of Brazilian Eupatorieae (Asteraceae) in the eastern cerrado and Atlantic forest. Molecular Phylogenetics and Evolution 97: 90-100.
  • Schilling, E. E. and D. E. Schilling. 2015. A Novel Hybrid Apomictic Eupatorium (Asteraceae) from the Southern Appalachians. Castanea 80: 273-277.
  • Schilling, E. E., R. W. Scott, and J. L. Panero. 2015. A revised infrageneric classification for Brickellia (Asteraceae, Eupatorieae). Phytotaxa 234: 151-158.
  • Schilling, E. E., A. F. Johnson, and G. D. Iacona. 2015. Coreopsis bakeri (Asteraceae; Coreopsideae), a new species from Florida, USA. Phytotaxa 231: 175-181.
  • Soto-Trejo, F., E. Schilling, S. Solórzano, K. Oyama, R. Lira, and P. Dávila. 2015. Phylogenetic relationships in the genus Florestina (Asteraceae, Bahieae). Plant Systematics and Evolution  1-14. DOI: 10.1007/s00606-015-1220-3
  • Schilling, E. E., A. Floden, and D. E. Schilling.  2015.  Barcoding the Asteraceae of Tennessee, tribe Cichorieae. Phytoneuron 2015-19: 1-8.
  • Schilling, E. E., J. L. Panero, B. S. Crozier, R. W. Scott, and P. Dávila.  2015.  Bricklebush (Brickellia) phylogeny reveals dimensions of the great Asteraceae radiation in Mexico.  Molecular Phylogenetics and Evolution 85: 161-170.
  • Schilling, E. E., N. Mattson, and A. Floden.  2014.  Barcoding the Asteraceae of Tennessee, tribe Coreopsideae. Phytoneuron 2014-34: 1-6.
  • Tippery, N. P., E. E. Schilling, J. L. Panero, D. H. Les, and C. S. Williams.  2014.  Independent origins of aquatic Eupatorieae (Asteraceae). Systematic Botany 39:1217-1225.
  • Elliott, A. G., C. Delay, H. Liu, Z. Phua, K. J. Rosengren, A. H. Benfield, J. L. Panero, M. L. Colgrave, A. S. Jayasena, K. M. Dunse, M. A. Anderson, E. E. Schilling, D. Ortiz-Barrientos, D. J. Craik, and J. S. Mylne.  2014.  Evolutionary origins of a bioactive peptide buried within preproalbumin. Plant Cell 26: 981-995.
  • Schilling, E. E., A. Floden, and S. B. Farmer. 2013. A new sessile-flowered Trillium species from Tennessee. Castanea. 78: 140-147.
  • Schilling, E. E. 2013. Barcoding the Asteraceae of Tennessee, tribes Cardueae and Vernonieae. Phytoneuron 2013-19: 1-7.
  • Schilling, E. E., J. L. Panero, B. S. Crozier, and P. Davila. 2013. Relationships of Asanthus (Asteraceae, Eupatorieae). Systematic Botany 38: 253-258.
  • Schilling, E. E. and A. Floden. 2012. Barcoding the Asteraceae of Tennessee, tribes Gnaphalieae and Inuleae. Phytoneuron 2012-99: 1-6.
  • Feild, T. S., D. S. Chatelet, L. Balun, E. E. Schilling, and R. Evans. 2012. The evolution of angiosperm lianescence without vessels - climbing mode and wood structure-function in Tasmannia cordata (Winteraceae). New Phytologist 193: 229-240.
  • Schilling, E. E. 2011. Systematics of the Eupatorium album complex (Asteraceae) from eastern North America. Systematic Botany 36: 1088-1100.
  • Schilling, E. E. and J. L. Panero. 2011. A revised classification of subtribe Helianthinae (Asteraceae: Heliantheae). II. Derived lineages. Botanical Journal of the Linnean Society 167: 311-331.
  • Schilling, E, E. 2011. Hybrid genera in Liatrinae (Asteraceae: Eupatorieae). Molecular Phylogenetics and Evolution, 59: 158-167.

Contact Information

  • 323 Hesler
  • Mailing Address: 569 Dabney Hall, 1416 Circle Dr, Knoxville, TN 37996
  • Phone: (865) 974-6200
  • Fax: (865) 974-3067
  • E-mail:
  • Lab Website