Edward Schilling


1978 – Ph.D., Indiana University

Research Interests

My research interests are in higher plant systematics with a focus on members of the sunflower family (Asteraceae).

Molecular Phylogenetic Analysis of Eupatorieae

A collaborative study being carried out together with Dr. Jose L. Panero of the University of Texas involves a systematic analysis of tribe Eupatorieae. This large group is primarily restricted to the New World, where it has produced an amazing amount of species-level diversity. Molecular phylogenetic analyses are being carried out with the goal of resolving issues of classification, and assessing the morphological changes associated with the success of this group.

Systematic Studies of Helianthus

The genus Helianthus comprises approximately 50 species native  to North America, and includes the agronomically important common sunflower, Helianthus annuus. Currently the focus of my studies are on the perennial species of the genus, with a particular emphasis on resolving issues of classification and working out the relationships of some of the polyploids, such as H. hirsutus, H. strumosus, and H. tuberosus.

Other Research in my Laboratory

Phylogenetic Analyses of Asteraceae of eastern North America – deciphering the relationships among species in the larger and more diverse genera of Asteraceae in eastern North America has proven to be challenging. Studies of Helianthus, Liatris, Eupatorium, and Solidago have revealed each to have a relatively low level of variability at the molecular level. We are seeking to find markers with an appropriate level of variability to help understand the causes for the contrasting high amount of species level diversity compared to the apparently low amount of genetic divergence.


  • Soto-Trejo, F., E. Schilling, S. Solórzano, K. Oyama, R. Lira, and P. Dávila. 2015. Phylogenetic relationships in the genus Florestina (Asteraceae, Bahieae). Plant Systematics and Evolution  1-14. DOI: 10.1007/s00606-015-1220-3
  • Schilling, E. E., A. Floden, and D. E. Schilling.  2015.  Barcoding the Asteraceae of Tennessee, tribe Cichorieae. Phytoneuron 2015-19: 1-8.
  • Schilling, E. E., J. L. Panero, B. S. Crozier, R. W. Scott, and P. Dávila.  2015.  Bricklebush (Brickellia) phylogeny reveals dimensions of the great Asteraceae radiation in Mexico.  Molecular Phylogenetics and Evolution 85: 161-170.
  • Schilling, E. E., N. Mattson, and A. Floden.  2014.  Barcoding the Asteraceae of Tennessee, tribe Coreopsideae. Phytoneuron 2014-34: 1-6.
  • Tippery, N. P., E. E. Schilling, J. L. Panero, D. H. Les, and C. S. Williams.  2014.  Independent origins of aquatic Eupatorieae (Asteraceae). Systematic Botany 39:1217-1225.
  • Elliott, A. G., C. Delay, H. Liu, Z. Phua, K. J. Rosengren, A. H. Benfield, J. L. Panero, M. L. Colgrave, A. S. Jayasena, K. M. Dunse, M. A. Anderson, E. E. Schilling, D. Ortiz-Barrientos, D. J. Craik, and J. S. Mylne.  2014.  Evolutionary origins of a bioactive peptide buried within preproalbumin. Plant Cell 26: 981-995.
  • Schilling, E. E., A. Floden, and S. B. Farmer. 2013. A new sessile-flowered Trillium species from Tennessee. Castanea. 78: 140-147.
  • Schilling, E. E. 2013. Barcoding the Asteraceae of Tennessee, tribes Cardueae and Vernonieae. Phytoneuron 2013-19: 1-7.
  • Schilling, E. E., J. L. Panero, B. S. Crozier, and P. Davila. 2013. Relationships of Asanthus (Asteraceae, Eupatorieae). Systematic Botany 38: 253-258.
  • Schilling, E. E. and A. Floden. 2012. Barcoding the Asteraceae of Tennessee, tribes Gnaphalieae and Inuleae. Phytoneuron 2012-99: 1-6.
  • Feild, T. S., D. S. Chatelet, L. Balun, E. E. Schilling, and R. Evans. 2012. The evolution of angiosperm lianescence without vessels – climbing mode and wood structure-function in Tasmannia cordata (Winteraceae). New Phytologist 193: 229-240.
  • Schilling, E. E. 2011. Systematics of the Eupatorium album complex (Asteraceae) from eastern North America. Systematic Botany 36: 1088-1100.
  • Schilling, E. E. and J. L. Panero. 2011. A revised classification of subtribe Helianthinae (Asteraceae: Heliantheae). II. Derived lineages. Botanical Journal of the Linnean Society 167: 311-331.
  • Schilling, E, E. 2011. Hybrid genera in Liatrinae (Asteraceae: Eupatorieae). Molecular Phylogenetics and Evolution, 59: 158-167.
  • Schilling, E, E. and J. L. Panero. 2010. Transfers to Simsia and description of Davilanthus, a new genus of Asteraceae (Heliantheae). Brittonia, 62: 309-320.
  • Ploch, S., Y.-J. Choi, C. Rost, H.-D. Shin, E. Schilling and M. Thines. 2010. Evolution of diversity in Albugo is driven by high host specificity and multiple speciation events on closely related Brassicaceae. Molecular Phylogenetics and Evolution 57: 812-820.
  • Robinson, H., E. Schilling, and J. L. Panero. 2009. Eupatorieae. Pp. 731-744 in Systematics, Evolution, and Biogeography of Compositae, eds. V. A. Funk, A. Susanna, T. F. Stuessy, and R. J. Bayer. Vienna: I.A.P.T.
  • Grubbs, K. C., R. L. Small, and E. E. Schilling. 2009. Evidence for multiple, autoploid origins of agamospermous populations in Eupatorium sessilifolium (Asteraceae). Plant Systematics and Evolution 279: 151-161.